Penn State fell to 3-4 -- and 0-4 in the Big Ten -- after blowing a late lead against Iowa in interim coach Terry Smith's first game since replacing James Franklin.

Penn State starting left guard Vega Ioane was listed as out for the Iowa game Saturday as the Nittany Lions lost, 25-24. The reason was undisclosed. Texas A&M transfer TJ Shanahan Jr. started in his place and handled all 76 snaps,

Penn State blew a fourth quarter lead. Iowa quarterback Mark Gronowski rushed for a career-high 130 yards and two touchdowns, including a 67-yard burst that set up Kaden Wetjen’s game-winning 8-yard touchdown run with 3:54 remaining.

With Saturday's loss to Iowa, Penn State fell under .500 for the first time since 2021, the same year it last lost four straight games.

Terry Smith has been named Penn State's interim football coach in the wake of James Franklin's firing. Here's what to know about the Nittany Lions' new leader. Now in his 12th year at Penn State, Smith entered this season for his fifth year as associate head coach of the Nittany Lions.

Terry Smith’s 3-4 Nittany Lions could not hold a 21-10 second-half lead in PSU’s 25-24 loss to Iowa on Saturday night at Kinnick Stadium.

Smith, who has served as the team's defensive recruiting coordinator since 2014, said he personally recruited three-fourths of the current roster. Now his job is keeping players from leaving.

Terry Smith says he's willing to do whatever he can to help Penn State win in his role as the interim head football coach of the Nittany Lions. Smith, who was named interim head coach of the team on Sunday following the firing of James Franklin, addressed the media on Monday at a press conference alongside athletic director Pat Kraft.

Penn State shocked the college football world this week when it parted ways with long-time head coach James Franklin, who had been with the program since 2014. Franklin led the Nittany Lions to a 104-45 (64-36) record over 12 seasons,